Missouri Department of Conservation

Taxonomy

Common Name: RABBIT, SWAMP

Phylum: CHORDATA

Class: MAMMALIA

Order: LAGOMORPHA

Family: LEPORIDAE

Genus: SYLVILAGUS

Species: AQUATICUS

SubSpecies: AQUATICUS

Taxonomic Authority: (BACHMAN)

Taxonomy References: 001

Status

GAME
STATE RANK S2
GLOBAL RANK G5

Status References: 027 , 003 , 006

Habitat Summary

"Occur in low forested wetlands along streams and drainage ditches. Prefer large forest tracts. Require mixture of forest size classes. Early successional stages are important."

Primary Habitat: "Forest - bottomland"

References: 002 , 007 , 008

Distribution

General Occurrence in State:

"Occurs in the southeast corner of the state *03,12,24*."

County Occurance

County Occurence
Known Likely Unknown Not Likely Historic Extirpated
BOLLINGER HOWELL ADAIR MCDONALD
BUTLER IRON ANDREW NEWTON
CAPE GIRARDEAU MADISON ATCHISON
CARTER OZARK AUDRAIN
DUNKLIN REYNOLDS BARRY
MISSISSIPPI BARTON
NEW MADRID BATES
OREGON BENTON
PEMISCOT BOONE
PERRY BUCHANAN
RIPLEY CALDWELL
SCOTT CALLAWAY
STE. GENEVIEVE CAMDEN
STODDARD CARROLL
WAYNE CASS
CEDAR
CHARITON
CHRISTIAN
CLARK
CLAY
CLINTON
COLE
COOPER
CRAWFORD
DADE
DALLAS
DAVIESS
DEKALB
DENT
DOUGLAS
FRANKLIN
GASCONADE
GENTRY
GREENE
GRUNDY
HARRISON
HENRY
HICKORY
HOLT
HOWARD
JACKSON
JASPER
JEFFERSON
JOHNSON
KNOX
LACLEDE
LAFAYETTE
LAWRENCE
LEWIS
LINCOLN
LINN
LIVINGSTON
MACON
MARIES
MARION
MERCER
MILLER
MONITEAU
MONROE
MONTGOMERY
MORGAN
NODAWAY
OSAGE
PETTIS
PHELPS
PIKE
PLATTE
POLK
PULASKI
PUTNAM
RALLS
RANDOLPH
RAY
SALINE
SCHUYLER
SCOTLAND
SHANNON
SHELBY
ST. CHARLES
ST. CLAIR
ST. FRANCOIS
ST. LOUIS
STONE
SULLIVAN
TANEY
TEXAS
VERNON
WARREN
WASHINGTON
WEBSTER
WORTH
WRIGHT

References for distribution: 012 , 020 , 024 , 025 , 003 , 034 , 008

Distribution by Watersheds

Castor R. and Castor R. Diversion Channel
Miss R. from Ohio R. to Arkansas Border
St. John's Ditch and Blue Ditch
St. Francis R. from Headwaters to Wappapello Dam
St. Francis R. from Wappapello Dam to Arkansas Border
Little R.
Area Bordered by Black R. and St. Francis R. South of Quintan, MO
Black R.
Current R.
Fourche Creek
Warm Fork Spring R. and South Fork
Eleven Point R.

Comments: "Likely to occur in units listed, based on county occurrence."

Distribution by Ecoregions

Mississippi River Alluvial Plains, Ozark Highlands

Comments: ""

Distribution by Potential Natural Vegetation

Oak-Hickory Forest
Oak-Hickory-Pine Forest
Southern Floodplain Forest

Distribution by Natural Divisions of Missouri

Ozark Border: Mississippi River
Ozark: St. Francois Mountains
Ozark: Lower Ozark
Mississippi Lowlands
Mississippi Lowlands: Crowley's Ridge
Mississippi Lowlands: Lowlands

Habitat Associations

Species is associated with "Terrestrial" habitats.

National Wetlands Inventory Association:

Palustrine
Riverine

Aquatic Associations:

"See Comments"
"Palustrine", "Emergent",
"Palustrine", "Emergent",
"Palustrine", "Forested, broad-leaved deciduous",
"Palustrine", "Forested, broad-leaved deciduous",
"Palustrine", "Forested, broad-leaved deciduous",
"Palustrine", "Forested, needle-leaved deciduous"
"Palustrine", "Scrub/shrub, broad-leaved deciduous",
"Palustrine", "Scrub/shrub, broad-leaved deciduous",
"Riverine, intermittent"
"Riverine, unknown perennial"

References for Aquatic Associations: 012 , 026 , 002 , 007 , 008

Habitat Types:

Wooded Riparian and Bottomland Hardwood
Swamp
Marsh
Permanent Stream

References for Habitat Types: 012 , 002 , 004 , 007 , 008

Terrestrial Natural Communities:

Forest
Bottomland Forest
Wet-Mesic Bottomland Forest
Wet Bottomland Forest
Flatwoods
Wetland
Marshes
Freshwater Marsh
Pond Marsh
Swamps
Swamp
Pond Swamp

References for Terrestrial Natural Communities: 012 , 002 , 004 , 007 , 008

Food Habits

Trophic Level:

"Herbivore"

Larval Food Habits

Comments for larval food habits:
""

References for larval food habits:

Juvenile Food Habits

Comments for Juvenile Food Habits:
""

References Juvenile Food Habits 008

Adult Food Habits

Cyperaceae (sedge); Bole/Stem
Lauraceae (laurel, sassafras); Bole/Stem
Rosaceae (rose, cherry, plum, blackberry); Bole/Stem
Anacardiaceae (cashew, sumac, poison ivy); Bole/Stem
Aquifoliaceae (holly); Bole/Stem
Liliaceae (lily, onion); Bole/Stem
Dicotyledoneae (dicots); Bark
Dicotyledoneae (dicots); Branches/Twigs
Lauraceae (laurel, sassafras); Branches/Twigs
Rosaceae (rose, cherry, plum, blackberry); Branches/Twigs
Aquifoliaceae (holly); Branches/Twigs
Cyperaceae (sedge); Leaves/Needles
Lauraceae (laurel, sassafras); Leaves/Needles
Rosaceae (rose, cherry, plum, blackberry); Leaves/Needles
Anacardiaceae (cashew, sumac, poison ivy); Leaves/Needles
Aquifoliaceae (holly); Leaves/Needles
Liliaceae (lily, onion); Leaves/Needles
Magnoliaceae (magnolia); Fruit/Seeds
Rosaceae (rose, cherry, plum, blackberry); Fruit/Seeds
Anacardiaceae (cashew, sumac, poison ivy); Fruit/Seeds
Aquifoliaceae (holly); Fruit/Seeds
Liliaceae (lily, onion); Fruit/Seeds
See comments on seasonal variation; Not Specified

Comments for Adult Food Habits:
""

References Adult Food Habits 014 , 018 , 019 , 002

Niche Requirements

Egg Niche Requirements

References for egg niches requirements:

Feeding Larvae Niche Requirements

References for feeding larvae niche requirements:

Resting Larvae Niche Requirements

References for resting Larvae niche requirements:

Feeding Juvenile Niche Requirements

Other niche requirements specified in comments(99999)
Floodplain
Inland wetlands: vegetated stream banks
Inland wetlands: swamp
Inland wetlands: marsh
Inland wetlands: slough, bayou
Inland wetlands: permanent stream
General habitat association specified in comments(00270)
Natural features: bottomland
Size of continuous forest stand specified in comments(00360)
Distance to water specified in comments(00430)
Canopy closure (%) specified in comments(00530)

References for feeding juvenile niche requirements: 013 , 018 , 002 , 007

Resting Juvenile Niche Requirements

Other niche requirements specified in comments(99999)
General habitat association specified in comments(00270)
Natural features: ridges
Size of continuous forest stand specified in comments(00360)
Distance to water specified in comments(00430)
Snags specified in comments(00510)
Canopy closure (%) specified in comments(00530)
Downed logs specified in comments(00660)
Stumps specified in comments(00720)

References for resting juvenile niche requirements: 013 , 018 , 002 , 007 , 011

Breeding Adult Niche Requirements

Other niche requirements specified in comments(99999)
Water level: semipermanently flooded
Water level: seasonally flooded
Water level: temporarily flooded
Floodplain
Inland wetlands: vegetated stream banks
Inland wetlands: swamp
Inland wetlands: marsh
Inland wetlands: slough, bayou
Inland wetlands: permanent stream
General habitat association specified in comments(00270)
Natural features: bottomland
Size of continuous forest stand specified in comments(00360)
Habitat size dependent on size of continuous wetland
Distance to water specified in comments(00430)
Leaf litter/ground debris specified in comments(00460)
Canopy closure (%) specified in comments(00530)
Shrubs specified in comments(00615)
Vines: greenbriar
Vines: honeysuckle
Vines specified in comments(00616)
Herbaceous cover (%) specified in comments(00620) Herbaceous cover (%) specified in comments
Stem density specified in comments(00650)
Downed logs specified in comments(00660)

References for feeding Adult niche requirements: 013 , 016 , 018 , 025 , 026 , 028 , 030 , 002 , 031 , 007 , 010

Feeding Adult Niche Requirements

Other niche requirements specified in comments(99999)
Aquatic vegetation: buttonbush (Cephalanthus)
Aquatic vegetation: willow (Salix)
Aquatic vegetation: reeds
Floodplain
Inland wetlands: vegetated stream banks
Inland wetlands: swamp
Inland wetlands: marsh
Inland wetlands: slough, bayou
Inland wetlands: permanent stream
General habitat association specified in comments(00270)
Natural features: bottomland
Size of continuous forest stand specified in comments(00360)
Distance to water specified in comments(00430)
Canopy closure (%) specified in comments(00530)
Shrubs: blackberry
Vines: poison ivy
Vines: grape
Vines: greenbriar
Vines: honeysuckle

References for feeding adult niche requirements: 013 , 018 , 028 , 002 , 007

Resting Adult Niche Requirements

Other niche requirements specified in comments(99999)
Aquatic vegetation: buttonbush (Cephalanthus)
Aquatic vegetation: willow (Salix)
Aquatic vegetation: reeds
Floodplain
Inland wetlands: vegetated stream banks
Inland wetlands: swamp
Inland wetlands: marsh
Inland wetlands: slough, bayou
Inland wetlands: permanent stream
General habitat association specified in comments(00270)
Natural features: bottomland
Size of continuous forest stand specified in comments(00360)
Distance to water specified in comments(00430)
Canopy closure (%) specified in comments(00530)
Shrubs: blackberry
Vines: poison ivy
Vines: grape
Vines: greenbriar
Vines: honeysuckle

References for resting adult niche requirements: 013 , 018 , 030 , 002 , 031 , 007 , 011

Niche Requirement Summary

Aquatic vegetation: buttonbush (Cephalanthus)
Aquatic vegetation: reeds
Aquatic vegetation: willow (Salix)
Brushpiles
Canopy closure (%) specified in comments(00530)
Distance to water specified in comments(00430)
Downed logs specified in comments(00660)
Floodplain
Forest openings (clearings, treefall gaps, etc.)
General habitat association specified in comments(00270)
Habitat size dependent on size of continuous wetland
Herbaceous cover (%) specified in comments(00620) Herbaceous cover (%) specified in comments
Inland wetlands specified in comments(00250)
Inland wetlands: marsh
Inland wetlands: permanent stream
Inland wetlands: slough, bayou
Inland wetlands: swamp
Inland wetlands: vegetated stream banks
Leaf litter/ground debris specified in comments(00460)
Natural features: bottomland
Natural features: ridges
Other niche requirements specified in comments(99999)
Shrubs specified in comments(00615)
Shrubs: blackberry
Size of continuous forest stand specified in comments(00360)
Snags specified in comments(00510)
Stem density specified in comments(00650)
Stumps specified in comments(00720)
Vines specified in comments(00616)
Vines: grape
Vines: greenbriar
Vines: honeysuckle
Vines: poison ivy
Water level: seasonally flooded
Water level: semipermanently flooded
Water level: temporarily flooded

Comments about Life History:

Code Comment
032 Population turnover estimated at 6.6 years *15*.
034 Habitat is limiting factor *07,13,02*.
007 In IN peak birth period: April and July *18*. TX breeding season nearly year-long with peaks in February and March *11*. In GA, breed March-September *13*. In MO, some breeding synchrony *02,17*. Females probably in estrus by early February in MO, with males attaining breeding condition earlier *19*. In a confined population, about 0.5 females pregnant by last week-February and remainder in March/early April, most final litters born late June through mid-July begin breeding early February *17*. Postpartum estrus *17,14,16*. Usually lasts less than1 hour *14*.
016 Nests usually bowl-shaped and placed in shallow excavation *10,16*. In a confined population, depression 4-7cm deep, mean length 15.5cm mean height 18.5cm *10*.
017 Gestation 39-40 days in TX *11*. 35-40 days in MO *02*. Two confined populations in MO found 37 days most common *10, 17*.
018 Pre-natal litter size in TX average 2.83 (1-5 young) *11*. Average 2.6 in GA *13*. Litter 1-6 in MO, with 3 young most frequently. *02,17*. Litters in miss. 1-3, average 2 *23*. Litter size estimate from corpus lutea counts average 3.7 (2-6), from embryo counts average 2.8 (1-4), from placental scar counts average 3.4 (3-4) *19*. Mean litter size 4.8 in confined MO population *10*.
019 Up to 5 litters annually *02*. May produce 4 litter/year *10*. Yearlings average more litters than adults- 3.5 to 2.8, respectively *17*.
022 No evidence of breeding in young-of-year *02,15*. Some females capable of breeding at 23-30 weeks old *17*.
028 In a MO study, females home range average 5.9 acre, (2.1 acre by minimum home range method), and male's averaged 4.6 acre (1.8 acre). Home range estimated by beagle chase: 4.1 acre-females, 2.0 acre-male *19*. In ge, average h. Range 19 acre *13*. In sw IL, home range of radio-tagged juveniles average .69 ha during flooding, .94 ha fall-winter without snow, .92 ha fall-winter with snow, home range of radio-tagged adults averaged: .64 ha fall-winter without snow, .52 ha fall-winter with snow, .53 ha during flooding. Home ranges of males were larger than those of females in spring-summer, but similar at other times. Some overlapping of home ranges occurred, but core areas were used almost exclusively *22*. In AR radiotelemetry study, average home range was 3.1 ha; large size of home range was due to rabbits moving onto levees or adjacent uplands during periods of flooding *30*.
035 Long periods of inundation increase nestling mortality *07,16*. Parasites include fleas, bot fly larvae, mites, ticks *02,11*. Predators include: hawks, owls, crows, foxes, coyotes, minks, weasels, snakes *02*.
099 Females may build 'dummy nests' before actual one built *02,16,10*. Swims and dives well, often uses tree roots and overhanging veg for protection *02*. Young leave nest during 2nd week, but cont. To nurse *02*. In a confined population, 76% young left nest at 10-15 days old *17*. A MO study showed total litter resorption may occur during severe floods, may be induced by adrenal stress syndrome *09*. In miss., 54% of population were juveniles *23*.

Life History Information

Home range size specified in comments(028)
Turnover rate specified in comments(032)
Limiting factors specified in comments(034)
Regulatory factors specified in comments(035)
Other life history information specified in comments(099)
Origin in state: native
Seasonal distribution in state: all seasons
Foraging strategy: grazing
Foraging strategy: browsing
Foraging sites: ground
Breeding season specified in comments(007)
Mating system: promiscuity
Duration of pair bond: no pair bond formed
Nest/den site: depression
Nest/den site: log
Nest materials: grasses
Nest materials: forbs
Nest materials: twigs
Nest materials: leaves
Nest materials: hair or feathers
Nest dimensions specified in comments(016)
Gestation/incubation period specified in comments(017)
Clutch/litter size specified in comments(018)
Number of broods/litters per year specified in comments(019)
Development of young at birth/hatching: altricial
Parental care of young: female
Age at sexual maturity specified in comments(022)
Sex ratio: 1:1

Comments about Life History:

Code Comment
032 Population turnover estimated at 6.6 years *15*.
034 Habitat is limiting factor *07,13,02*.
007 In IN peak birth period: April and July *18*. TX breeding season nearly year-long with peaks in February and March *11*. In GA, breed March-September *13*. In MO, some breeding synchrony *02,17*. Females probably in estrus by early February in MO, with males attaining breeding condition earlier *19*. In a confined population, about 0.5 females pregnant by last week-February and remainder in March/early April, most final litters born late June through mid-July begin breeding early February *17*. Postpartum estrus *17,14,16*. Usually lasts less than1 hour *14*.
016 Nests usually bowl-shaped and placed in shallow excavation *10,16*. In a confined population, depression 4-7cm deep, mean length 15.5cm mean height 18.5cm *10*.
017 Gestation 39-40 days in TX *11*. 35-40 days in MO *02*. Two confined populations in MO found 37 days most common *10, 17*.
018 Pre-natal litter size in TX average 2.83 (1-5 young) *11*. Average 2.6 in GA *13*. Litter 1-6 in MO, with 3 young most frequently. *02,17*. Litters in miss. 1-3, average 2 *23*. Litter size estimate from corpus lutea counts average 3.7 (2-6), from embryo counts average 2.8 (1-4), from placental scar counts average 3.4 (3-4) *19*. Mean litter size 4.8 in confined MO population *10*.
019 Up to 5 litters annually *02*. May produce 4 litter/year *10*. Yearlings average more litters than adults- 3.5 to 2.8, respectively *17*.
022 No evidence of breeding in young-of-year *02,15*. Some females capable of breeding at 23-30 weeks old *17*.
028 In a MO study, females home range average 5.9 acre, (2.1 acre by minimum home range method), and male's averaged 4.6 acre (1.8 acre). Home range estimated by beagle chase: 4.1 acre-females, 2.0 acre-male *19*. In ge, average h. Range 19 acre *13*. In sw IL, home range of radio-tagged juveniles average .69 ha during flooding, .94 ha fall-winter without snow, .92 ha fall-winter with snow, home range of radio-tagged adults averaged: .64 ha fall-winter without snow, .52 ha fall-winter with snow, .53 ha during flooding. Home ranges of males were larger than those of females in spring-summer, but similar at other times. Some overlapping of home ranges occurred, but core areas were used almost exclusively *22*. In AR radiotelemetry study, average home range was 3.1 ha; large size of home range was due to rabbits moving onto levees or adjacent uplands during periods of flooding *30*.
035 Long periods of inundation increase nestling mortality *07,16*. Parasites include fleas, bot fly larvae, mites, ticks *02,11*. Predators include: hawks, owls, crows, foxes, coyotes, minks, weasels, snakes *02*.
099 Females may build 'dummy nests' before actual one built *02,16,10*. Swims and dives well, often uses tree roots and overhanging veg for protection *02*. Young leave nest during 2nd week, but cont. To nurse *02*. In a confined population, 76% young left nest at 10-15 days old *17*. A MO study showed total litter resorption may occur during severe floods, may be induced by adrenal stress syndrome *09*. In miss., 54% of population were juveniles *23*.

References for life history: 013 , 014 , 015 , 016 , 017 , 018 , 019 , 022 , 023 , 002 , 008 , 009 , 010 , 011

Management

Beneficial Management Practices:

Regulate harvest of species being described
Water - develop/maintain wetlands
Water - control pollution (thermal, physical, chemical)
Water - develop/maintain streamside vegetation
Water - supplemental feeding (winter, spring, etc.)
Forest - develop and maintain food plots
Forest - maintain riparian habitats
Forest - maintain streamside vegetation
Forest - maintain habitat diversity
Forest - uneven age timber management
Forest - reforestation
Forest - develop/maintain brushpiles

Beneficial Management References: 021 , 002 , 007 , 008

Adverse Managment Practices:

Water - dredging and filling
Water - navigational improvements (channelization, dams, locks)
Water - stream channelization
Water - drainage of wetlands, marshes, ponds, lakes
Forest - even age timber management

Adverse Management References: 002 , 007 , 008

Comments on Management:
Conversion of bottomland forests to agriculture has led to a decline in numbers and a more restricted distribution *29*. May increase distribution by stocking along selected sites on Ozark streams *07*. Habitat is enhanced by disturbance *31*. Density of rabbits may be increased by creating canopy openings in forest. Suggest clearings of 0.1-0.5 ha spaced several hundred meters apart in even-age forests, this needs more research however *07*. Selective cutting to open the forest canopy and promote growth of early successional ground cover is beneficial. Should leave slash piles created during cutting to enhance winter habitat and survival. it is also beneficial to improve habitat on areas where rabbits seek refuge during extended periods of flooding. *28*. Riparian corridors should be protected from grazing and habitat fragmentation *31*.

References for Management Comments: 028 , 029 , 031 , 007 , 008

References

Reference Code Citation
001 Hall, E.R. 1981. The Mammals Of North America, 2nd Ed. John Wiley And Sons, Inc. New York. 1181pp.
002 Schwartz, C.W. and E.R. Schwartz. 1981. The Wild Mammals Of Missouri 2nd Ed. Univ. Of Missouri Press And Mo Dept. Of Conservation, Columbia, Mo. 356 Pp.
003 The Checklist Of Rare And Endangered Species Of Missouri. 1991. Missouri Department Of Conservation. Jefferson City, Mo. 44 Pp.
004 Kelly, G. (Ed.) 1986. Animal Habitat Relations Handbook. Mo Dept. Of Conservation and U.S.D.A. Forest Service. Jefferson City, Mo. 293 Pp.
005 Thom, R.H. and J.H. Wilson. 1980 The Natural Divisions Of Missouri. Trans. Mo Acad. Sci. 14:9-24.
006 The Wildlife Code of Missouri. Missouri Department of Conservation, P.O. Box 180, Jefferson City, MO 65102. 573-751-4115.
007 Unpb Korte, P.A. 1975. Distribution And Habitat Requirements Of The Swamp Rabbit In Missouri. M.S. Thesis. Univ. Mo, Columbia. 127 Pp. (112 Stephens Hall, Columbia, Mo 65211 (573)882-3436.)
008 Unpb Sadler, K.C. Mo Dept. Conserv. 1110 S. College Ave. Columbia, Mo. 65201. (573)882-9880.
009 Conaway, C.H., T.S. Baskett, and J.E. Toll. 1960. Embryo Resorption In The Swamp Rabbit. J. Wildl. Manage. 24(2):197-202.
010 Holler, N.R., T.S. Baskett, and J.P. Rogers. 1963. Reproduction In Confined Swamp Rabbits. J. Wildl. Manage. 27(2):179-183.
011 Hunt, T.P. 1959. Breeding Habits Of The Swamp Rabbit With Notes On Its Life History. J. Mammal. 40(1):82-91.
012 Korte, P.A. and L.H. Frederickson. 1977. Swamp Rabbit Distribution In Missouri. Trans. Mo Acad. Sci. 10-11:72-77.
013 Lowe, C.E. 1958. Ecology Of The Swamp Rabbit In Georgia. J. Mammal. 39(1):116-127.
014 Marsden, H.M. and N.R. Holler. 1964. Social Behavior In Confined Populations Of The Cottontail And The Swamp Rabbit. Wildl. Monogr. 13. 39pp.
015 Martinson, R.K., J.W. Holten, and G.K. Brakhage. 1961. Age Criteria And Population Dynamics Of The Swamp Rabbit In Missouri. J. Wildl. Manage. 25(3):271-281.
016 Sorensen, M.F., J.P. Rogers, and T.S. Baskett. 1972. Parental Behavior In Swamp Rabbits. J. Mammal. 53(4):840-849.
017 Sorensen, M.F., J.P. Rogers, and T.S. Baskett. 1968. Reproduction And Development In Confined Swamp Rabbits. J. Wildl. Manage. 32(3): 520-531.
018 Terrel, T.L. 1972. The Swamp Rabbit (Sylvilagus Aquaticus) In Indiana. Am. Midl. Nat. 87(2):283-295.
019 Toll, J.E., T.S. Baskett, and C.H. Conaway. 1960. Home Range, Reproduction, And Foods Of The Swamp Rabbit In Missouri. Am. Midl. Nat. 63(2):398-412.
020 Unpb Elder, W.H. 112 Stephens Hall, Umc, Columbia, Mo 65211. (573) 882-3436.
021 Missouri Dept. Of Conservation and U.S. Forest Service. 1985. Management Of Snags And Cavity Trees In Missouri. Mo Cons. Comm, Jefferson City, Mo. 21 Pp.
022 Kjolhaug, M.S. and A. Woolf. 1988. Home Range Of The Swamp Rabbit In Southern Illinois. J. Mammal. 69(1):194-197.
023 Palmer, W.E., G.A. Hurst, B.D. Leopold and D.C. Cotton. 1991. Body Weights And Sex And Age Ratios For The Swamp Rabbit In Mississippi. J. Mammal. 72:620-622.
024 Unpb Missouri Department of Conservation Heritage Database. P.O. Box 180, Jefferson City, Mo 65102.
025 Unpb Dailey, T.V. 1992. Swamp Rabbit Distribution Study. Mo Dept. Of Conservation P-R Project W-13-R-46, Study No. 46, Job No. 2. 75 Pp.
026 Sole, J.D. 1994. Assessing Swamp Rabbit Distribution In Kentucky. Proc. Ann. Conf. Southeast Assoc. Fish Wildl. Agencies 48:145-151.
027 Missouri Natural Heritage Program. 2004. Missouri Species and Communities of Conservation Concern Checklist. Missouri Department Of Conservation. Jefferson City, Missouri. 47 Pp.
028 Unpb Porath, J.W. 1997. Swamp Rabbit (Sylvilagus Aquaticus) Status, Distribution, And Habitat Characteristics In Southern Illinois. M.S. Thesis, S. Il. Univ., Carbondale. 61 Pp.
029 Woolf, A. and M. Barbour. 1998. Illinois Swamp Rabbit Study. Il. Dept. Nat. Res., Div. Of Wildl. Resources. P-R Project W-127-R-3. 90 Pp.
030 Zollner, P.A., W.P. Smith and L.A. Brennan. 2000. Home Range Use By Swamp Rabbits (Sylvilagus Aquaticus) In A Frequently Inundated Bottomland Forest. Amer. Midl. Nat. 143(1):64-69.
031 Baccus, J.T. and M.W. Wallace. 1997. Distribution And Habitat Affinity Of The Swamp Rabbit (Sylvilagus Aquaticus: Lagomorpha: Leporidae) On The Edwards Plateau Of Texas. Occas. Papers, Mus. Texas Tech. Univ. No. 167. 16 Pp.
033 Scheibe JS and R Henson. 2003. The distribution of swamp rabbits in southeast Missouri. Southeastern Naturalist 2(3): 327-334.
034 UNPB Shelby Timm. MDC Mammal Database. Missouri Dept. of Conservation, Ellington Office. 2929 County Rd 618, Ellington, MO 63638/